Parents

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Asa Gray post-doc 1872 Harvard

Children

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William T. Hornaday research assistant 1872-1873 Iowa State (Neurotree)
Cornelius Lott Shear research assistant 1897 University of Nebraska - Lincoln (Plant Biology Tree)
Herbert Osborn grad student 1880 Iowa State (Evolution Tree)
Conway Macmillan grad student 1888 University of Nebraska (Plant Biology Tree)
Roscoe Pound grad student 1897 University of Nebraska
Frederic E. Clements grad student 1898 University of Nebraska
John E. Weaver grad student 1908 University of Nebraska
Raymond John Pool grad student 1913 University of Nebraska
Melvin Randolph Gilmore grad student 1914 University of Nebraska (Plant Biology Tree)
Paul Bigelow Sears grad student 1915 University of Nebraska
Theodore Alexander Kiesselbach grad student 1918 University of Nebraska - Lincoln (Plant Biology Tree)
Joseph ​C Arthur grad student 1870-1973 Iowa State College (Evolution Tree)
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