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Jan Willem van Wijhe research assistant 1926 RUG
Martinus Willem Woerdeman grad student 1928 RUG
 (Over het sluiten van de neuraalbuis : een experimenteel-morphologische studie bij urodelen)


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Pieter Jacob Donker grad student 1951 Amsterdam (Physiology Academic Tree)
Maarten Vink grad student 1952 Amsterdam (Physiology Academic Tree)
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